Research on soldier ants reveals that evolution can go in reverse
https://phys.org/new...on-reverse.html
"The shape and size of a turtle-ant soldier's head is dictated by the type of tunnel the species in question occupies. The ants don't dig the tunnels themselves, but move into those excavated by wood-boring beetles. And since a hand-me-down tunnel might be too big or too small, Kronauer says, the ants diversify rapidly to be able to occupy it."
"In some cases, more specialist species reversed direction over time, evolving back into more generalist head shapes."
Past & Present
Veromessor pergandei, andrei, stoddardi; Novomessor cockerelli
Camponotus fragilis, Camponotus sansabeanus (inactive), vicinus, laevigatus/quercicola, CA02
Pogonomyrmex subnitidus, P. californicus (inactive)
Liometopum occidentale (inactive); Prenolepis imparis; Myrmecocystus mexicanus (inactive); Tetramorium sp. (inactive); Lasius sp.
Termites: Zootermopsis angusticollis, and a box of drywood termites that can't be seen
Isopods: (most no longer keeping) A. gestroi, granulatum, kluugi, maculatum, vulgare; C. murina; P. hoffmannseggi, P. haasi, P. ornatus; V. parvus, P. pruinosus, T. tomentosa
Spoods: (no longer keeping) Phidippus sp., other